Ichika Usami is a second year middle school student whose family runs the Kirakira Patisserie shop. One day, she encounters a fairy named Pekorin and learns that evil monsters known as Henteko have been stealing "Kirakiraru", the energy that resides within sweets and desserts. Thus, Ichika becomes a Pretty Cure and, alongside four other cures, forms the Kirakira PreCure a la Mode to fight off the Henteko and protect the sweets.

The year is 1863, with the story being based in Kyoto. We follow a youth named Nio who is an honest and kind soul who's considerate of his family, yet he also has a hidden burning passion for seeking justice. From the time he came across Toshizou Hijikata and Souji Okita, members from a group of hated ronin known as the Miburo, his life then takes a turn to a bluer, and clearer path. Nio, alongside the other men whose hearts burn bravely and true, step forth towards an honest path with their blue wills adorned on their hearts!

Caught in a 150-year-long war, interstellar nations reach the pinnacle of strategic combat at the hands of two genius leaders. Reinhard of the Galactic Empire, and Yang of the Free Planets Alliance lead the charge from opposing fronts. Fighting is their destiny, but in this vast universe torn by political intrigue, their greatest enemy may not be each other.