年轻男子雅各布相信自己是一只困在人体内的狼，吃住行都像狼一样，这是一种被称为“物种焦虑症”的现实状况，他被送到一家诊所接受治疗，遇到了其他有同样情况的病人，他们被迫在冷酷无情的医生“动物园管理员”的手中接受极端治疗。当雅各布遇到神秘的“野猫”后，两人的友谊发展成了不可否认的迷恋，雅各布面临一个难题：他是否愿意为了爱情放弃真我？

两个童年时期在寄养系统中经历了重重波折的疏远兄弟，突然间不得不共同生活在一个屋檐下。他们面临着重新审视过去、与过去的自己和解的挑战，同时也要共同思考和决定未来的方向。

Joseph falls into despair when his nine-year-old son Shea leaves for Australia with his ex Debbie. Suffering the hangover from hell, he walks away from his present life and boards a boat bound for Ireland to confront memories from his childhood.

In 1985, while working as a coal merchant to support his family, Bill Furlong discovers disturbing secrets kept by the local convent and uncovers truths of his own; forcing him to confront his past and the complicit silence of a small Irish town controlled by the Catholic Church.

Set during Christmas 1988, Lol is haunted by the devastating events that took place two and a half years before. She and Woody both find themselves struggling to cope with their lives without each other after he leaves the gang. Lol is carrying the burden of her guilt, whilst Woody is trying to build a domestic life with a new girlfriend and a potential promotion at work. Shaun has started drama college and, although still in a relationship with Smell, he has grown close to a girl performing in his Christmas play.

Shaun, Harvey, Gadget, Trev and Kelly hit up the nightlife of raves and ecstasy. Woody and Lol are happy, living together with their kids and Combo is still in prison. But things slowly change. This is the year 1990 and This is England.

Maud's best friend Elizabeth has disappeared, but as she tries to solve the mystery, dementia threatens to erase all the clues, giving the search a poignant urgency.