Yukihira Souma's dream is to become a full-time chef in his father's restaurant and surpass his father's culinary skill. But just as Yukihira graduates from middle schools his father, Yukihira Jouichirou, closes down the restaurant to travel and cook around the world. Although downtrodden, Souma's fighting spirit is rekindled by a challenge from Jouichirou which is to survive in an elite culinary school where only 10% of the students graduate. Can Souma survive?

Famous racehorses that have left behind worthy legacies, unique as they can be, are reincarnated as horse girls in a parallel world. In this life, they start their journey anew as they continue to race and perhaps relive the success they once lived through. Aspiring to become the best racehorse in Japan, a horse girl named Special Week moves to Tokyo to enroll in the Tracen Academy—an institution that nurtures horse girls like her to become better racers. There, Special Week witnesses the sophisticated running style of Silence Suzuka and is inspired to become a racer like her. Shortly after, Special Week finds herself recruited into Silence Suzuka's team, Spica. From there, she begins her path to the top—one lap at a time.

Ten years have passed since the end of Bakumatsu, an era of war that saw the uprising of citizens against the Tokugawa shogunate. The revolutionaries wanted to create a time of peace, and a thriving country free from oppression. The new age of Meiji has come, but peace has not yet been achieved. Swords are banned but people are still murdered in the streets. Orphans of war veterans are left with nowhere to go, while the government seems content to just line their pockets with money.