Constantly outperformed by the girls' club, the boys' soft tennis club faces disbandment due to their poor skills and lack of positive results. In desperate need of capable players, Toma Shinjou is looking to recruit, but he fails to do so. Enter Maki Katsuragi, a transfer student who demonstrates great reflexes when he catches a stray cat, Instantly capturing Toma's attention. Toma ambitiously asks Maki to join the team but is quickly rejected, as Maki doesn't wish to join any clubs. Toma refuses to back down and ends up persuading Maki - only under the condition that Toma will pay him for his club expenses.

As Maki joins the team, he immediately outshine the rest of the team. This gives rise to conflict among the team, Maki challenges his fellow team members to devote themselves to the game they once neglected.

This story focuses on the boys' soft tennis club and their discovery of their own capability, while also enduring the darker side of growing up in middle school.

While W-0 is away, Leila and the remaining crew at base find themselves surrounded by Britannian Knightmares. Piloting the Vercingetorix, Shin Hyuuga leads the enemy charge, having usurped the position of Grand Master in the Knights of St. Michael. With her core members unavailable, it's up to Leila to protect her comrades, but this may be much tougher than she realizes: a spy lurking among them is finally ready to reveal himself. Left with no choice but to surrender, Leila arranges a meeting with Shin in order to ensure the safety of her squad. But as negotiations begin to fall apart, Akito and an unexpected ally return in the nick of time to turn things around.

The Ark Fleet has been destroyed, and a significant number of the enemy's troops have been wiped out due to its crash landing. As the remaining forces of the Holy Order of Michael regroup in order to launch a final assault on Weiswolf Castle, the wZERO unit, along with their new ally Ashley Ashra, stand ready to intercept them. Meanwhile, with his Geass out of control, Shin moves to erase his younger brother's existence once and for all. But Akito, having promised Leila that he will come back alive, refuses to accept such a fate, and the two clash in their final battle.