High schoolers Mitsuha and Taki are complete strangers living separate lives. But one night, they suddenly switch places. Mitsuha wakes up in Taki’s body, and he in hers. This bizarre occurrence continues to happen randomly, and the two must adjust their lives around each other.

宇宙中的一场爆炸让地球上的熊类发生了暴走，身强体壮的熊类很快就让人类伤亡惨重，一场旷日持久的人熊之战就此拉开序幕。为了生存，人类建起了高耸的墙壁，将熊隔离在了墙壁之外。然而，他们并不知道的是，在这个世界上，危险的并不仅仅是那些凶猛强壮的野兽们。
岚丘学园学院中，来了两名转校生，她们是百合城银子（荒川美穂 配音）和百合咲露露（生田善子 配音），表面上看来，她们只是柔软可爱的女孩子，然而，两人的真实身份，竟然是可以化身成为人类的熊。外冷内热的椿辉红羽（山根希美 配音）、外柔内刚的泉乃纯花（小仓唯 配音）、善良开朗的百合园蜜子（悠木碧 配音），在百合城银子和百合咲露露的眼中，她们都是散发着香味的猎物。

Humans have created a Wall of Severance to separate themselves from the bears, who grew violent and attacked humans after a far-off planet known as Kumalia exploded many years ago, turning into a meteor shower that fell upon earth. Two bears, Ginko Yurishiro and Lulu Yurigasaki, sneak through the Wall of Severance and disguise themselves as humans, enrolling in Arashigaoka Academy and taking an interest in Kureha Tsubaki, a human girl who despises bears.

曾经，来自海洋的神秘敌人“塞壬”向人类发起了猛烈的进攻，“碧蓝航线”的少女们凭借着她们钢铁般的意志与敌人奋勇作战，最终击败了对手，守护了人类。然而，随着时间的流逝，碧蓝航线内部因为理念的差异逐渐产生分化，衍生出了名为“赤色中轴”大的对立阵营。
被称为“灰色的幽灵”的最强英雄企业（石川由依 配音）、每天都迷迷糊糊但关键时刻意外的顶用的拉菲（长绳麻理亚 配音）、虽然年纪不大但成熟体贴的大黄蜂（山本希望 配音），个性好强但心地善良的哈曼（伊藤明日香 配音），面对动荡的时局，当这些少女们再一次被推上战场，会发生怎样的故事呢？

When the "Sirens," an alien force with an arsenal far surpassing the limits of current technology, suddenly appeared, a divided humanity stood in complete solidarity for the first time. Four countries formed Azur Lane, paving the way for the improvement of modern warfare, which led to an initial victory. However, this tenuous union was threatened by opposing ideals, dividing the alliance into two and humanity became fragmented once again.

As a seasoned and experienced fighter, the "Grey Ghost" Enterprise shoulders Azur Lane's hope for ending the war. But behind her stoic persona hides a frail girl, afraid of the ocean. Even so, she continues to fight as she believes that it's the only purpose for her existence.

Amidst the neverending conflict within humankind, the keys that could unite a fragmented race might exist: a soldier coming to terms with her mysterious personality and camaraderie between those with different ideals.

In the fallout of nuclear war, the last of humanity survives with the help of life-like androids known as Dolls. Built to serve, some fight in wars as Tactical Dolls or T-Dolls. Now, an elite team comprised of T-Dolls are sent to face a new rogue threat, Sangvis Ferri. Leading them into battle is their new human commander, M4A1.

大场奈奈。
圣翔音乐学园第99期生 学号15号。
中学时代孤身一人身处演剧部的她，知晓了在这所学园首次创设舞台的真正喜悦。
以同一个舞台为目标，携手共进的99期生这些同志。
落幕的瞬间沸腾的掌声，以及令身体颤抖的充实。
让那如同燃烧宝石般的光辉
让那如同闪亮彩虹般的每个幸福之日，再一次——
被那份眩目囚禁的奈奈，参加了长颈鹿主办的谜之选拔。
成为合格者的她，期盼着在命运舞台上的「第99回圣翔祭，那一年间的再演」。
「好吧」
然后再次开始的，「那一年」。
奈奈将开花结果的才能，全部投入到那眩目之日的「再演」中。
但她尚未知晓。
没有完全相同的舞台。
没有不会结束的舞台。
在重复的再演前方，旋转的轮舞开幕。
「本该是演出过的舞台。但，却是我不知道的舞台」
「那就是，轮舞·轮舞·轮舞——我明白的」

After the Third World War in 206x AD, the world has been devastated, the tactical dolls of the private military company Griffin are in an endless battle with the “Iron-Blood Construction Company,” which deceived mankind.