152 years after the events of season two, The Foundation has become increasingly established far beyond its humble beginnings while the Cleonic Dynasty's Empire has dwindled. As both of these galactic powers forge an uneasy alliance, a threat to the entire galaxy appears in the fearsome form of a warlord known as "The Mule," whose sights are set on ruling the universe by use of physical and military force as well as mind control. It’s anyone's guess who will win, who will lose, who will live and who will die as Hari Seldon, Gaal Dornick, the Cleons and Demerzel play a potentially deadly game of intergalactic chess.

When the initial Cylon attack against the Twelve Colonies fails to achieve complete extermination of human life as planned, twin Number Ones (Cavils) embedded on Galactica and Caprica must improvise to destroy the human survivors.

As season 4 begins, the last remnants of humanity continue their search for a new home. Galactica’s crew attempts to make sense of the inexplicable while four members of the fleet are still reeling from past revelations.

In the year 2517, after the arrival of humans in a new star system, follow the adventures of the renegade crew of Serenity, a "Firefly-class" spaceship.

More than a century after the events in the Season 1 finale, tension mounts throughout the galaxy. As the ruling dynasty begins to unravel, a young queen plots her revenge. Meanwhile, a colony of people with special abilities poses a new threat. And it’s all leading toward a collision course for war.