Interpol Inspector Zenigata begins a world-wide hunt for the infamous thief Arsène Lupin III, who, with his cavalcade of collaborators infiltrate a highly secured fortress-like safe named the 'Ark of Navarone' to claim the "Crimson Heart of Cleopatra".

Young assassins Azumi and Nagara continue their mission to prevent a civil war. In their hunt for Masayuki Sanada, who is protected by both an army and a dangerous clan, they meet Ginkaku, a person who shows a remarking resemblance with former friend Nachi.

Genji and his victorious G.P.S. alliance find themselves facing down a new challenge by the students of Hosen Academy, feared by everyone as 'The Army of Killers.' The two schools, in fact, have a history of bad blood between them. And the simmering embers of hatred are about to flare up again, burning away any last remnants of the truce they had so rigorously observed until now.