亚里安（中原茂 配音）本是普罗米修斯和潘多拉的孩子，然而潘多拉在还未生下亚里安之时就战死了，普罗米修斯拼尽全力才让亚里安有了降临这个世界的机会。之后，普罗米修斯因为盗取圣火而遭到了宙斯的处罚，亚里安辗转被德墨忒尔收养，可很快，德墨忒尔也遭遇了悲惨的命运，冥王哈迪斯趁机掳走了亚里安，令他相信自己是海王波塞冬的儿子，为了寻找父亲，亚里安踏上了旅途，不仅如此，他还发誓要杀死宙斯为自己的亲人们复仇。
诸神之战开始了，雅典娜作为宙斯一方，带领着地上的人民和海王展开了殊死搏斗，而亚里安则彻底沦为哈迪斯利用的工具。

西元2003年5月，金星相隔3千年后再度与巨大冰状行星冲撞，厚实的碳酸大气层因此遭到破坏，接着地表开始产生酸性海洋，并出现了地球型的大气层。人类随即开始着手金星的地球化以及移民计划，并于2018年（金星历元年）将首批移民团送至伊修塔尔大陆，但由于金星的土地却超乎想像的贫瘠，于是人们又再度为了争夺土地而掀起战端。金星历72年，金星被分割成伊修塔尔与爱芙罗黛蒂两大自治州，第二、三批移民团也相继抵达，独立通信社记者，苏珊．索玛兹亦是其中之一。此时，让金星的人们最感到热衷的就是在爱芙罗黛蒂的首都．伊娥市所进行的机车大赛“绕圈赛”；参赛车队之一的“杀手突击队”在蜜兰达的领导下，希洛、玛姬等人也都有相当优异的表现。某日在比赛的途中，突然出现了数百辆的伊修塔尔重型战车“章鱼”，伊娥市因而遭到占领，希洛等人也遭受到了迫害，后来“杀手突击队”的这群年轻人终于挺身而出，试图抵抗这群侵略者。拯救了希洛的爱芙罗黛蒂军游击队“猎犬”队长．卡兹上尉，送给了希洛一部特殊机车武器“猎犬”。在不安与绝望之中，希洛、玛姬、蜜兰达等人也接连地被卷入这场战争……

Universal Century 0079. Humanity has turned even space itself into a battlefield, and the Principality of Zeon forces advance after wiping out half the world's population by carrying out the Operation British (colony drop). In response, the Earth Federation Forces mobilize their overwhelming fighting strength to regain the advantage. The complex intrigues of the Zabi family... Sayla Mass, as she contends with the whims of fate... Hamon and Ramba Ral, who is now a pilot in the Zeon forces... Amuro and Fraw, leading peaceful lives at Side 7... a dark shadow falls across them all. And the Zeon ace Char Aznable, driven by revenge, goes into action as the "Battle of Loum" finally begins.

Universal Century 0068, Side 3 - The Autonomous Republic of Munzo. Zeon Zum Deikun attempts to declare complete independence of Munzo from the Earth Federation Government, while he preaches the evolutionary potential of humans who have advanced into outer space. Deikun however suddenly falls to his death in the middle of his speech at parliament. Upon Deikun's death, Jimba Ral spreads word of a House of Zabi conspiracy - but despite such efforts the power and sphere of influence of House of Zabi, led by Degwin Sodo Zabi, only seem to escalate. While we witness for the first time the untold convulsions of Universal Century history, Casval and Artesia, just bereaved of their father, must face destinies which will be just as tumultuous as the very era itself.

Enter the tale of the Crusher Council, a group of rugged individuals known for assignments ranging from transportation to terraforming and everything in between. In the early days of space exploration the Crushers took on the job of destroying asteroids and defining space lanes. Because of their work, they were nicknamed "Crushers" which eventually became their business moniker. Despite the rough and ready nature of the Crushers' work, they subscribe to a few steadfast rules. Unethical and illegal assignments are taboo, and any Crusher accepting one is barred from the Union. Of course, this presents problems for shady clients who try to trick the Crushers into accepting misleading assignments. They know that once the Union accepts a case the Crushers are honor-bound to follow it through. Among the various worlds, the Crusher Council has a stunning reputation, and among the Crushers, the most elite team is the one led by Crusher Dan and his successor, Crusher Joe.